Depositional Controls on the Ichnology of Ordovician Wave-dominated Marine Facies：New Evidence from the Shirgesht Formation,Central Iran

The Lower Ordovician Shirgesht Formation in central Iran is composed of siliciclastic and carbonate rocks deposited in diverse coastal and marine shelfal environments（tidal flat,lagoon,shoreface,offshore-shelf and carbonate ramp）.Five facies associations contain diverse ichnofossil assemblages that show distinct proximal to distal trends formed in a wide range of physical-chemical conditions.The ethological groups of trace fossils in the Shirgesht Formation reflect a gradient of depositional stress conditions across a wave-influenced shoreline and shelf.Deposits of waveinfluenced environments make up a significant component of the geological record of shallow marine settings,and the ability to determine paleoenvironments in detail in such successions is critical for reconstruction of depositional histories and sequence-stratigraphic interpretation.The Cruziana ichnofacies of the study shows highly diverse suites that record the establishment of a benthic community under stable conditions and a long-term colonization window.The Skolithos ichnofacies recognized is a low diversity opportunistic ichnocommunity suite that resulted from colonization after tempestite deposition in a stressed environment.The strata record an onshore to offshore replacement of the Cruziana ichnofacies（with abundant feeding traces of deposit-feeders） by the Skolithos ichnofacies（dominated by suspension-feeders and predators）.A transitional zone between the two ichnofacies coincides with the offshore-transition/distal lower-surface deposits.The distribution of ichnofacies,the diversity and range of ethological characteristics reflected by the ichnogenera,and the wide range of wave-dominated coastal facies demonstrate the potential to use individual trace fossils and ichnofacies for significantly refined palaeoenvironmental analysis of wavedominated coastal settings,particularly in Ordovician successions.