On the systematics of Anania Huebner, 1823 (Pyraloidea: Crambidae: Pyraustinae)

Currently, Pyraustinae (Lepidoptera: Pyraloidea: Crambidae) are split into many genera that often contain a small number of species only. This classification is largely influenced by traditional and typological concepts and do not necessarily reflect natural relationships. Thus, we encourage the idea to fuse taxa based on synapomorphies, as suggested by Leraut (2005), who argued, that an elongated, serrated sclerite of the phallus in males and a digitiform structure freely extending into the antrum in females is apomorphic for members of Anania Huebner, 1823. Screening the literature, we found four further species belonging to this monophylum: Anania hasanensis (Kirpichnikova, 1998) (Opsibotys) comb. n., Anania luteorubralis (Caradja, 1916) (Pyrausta) comb. n., Anania obtusalis (Yamanaka, 1987) (Perinephela) comb. n., and Anania shafferi (Speidel & Hanigk, 1990) (Algedonia) comb. n. Investigating Chinese Pyraustinae, we also found these characters in taxa which so far were not assigned to Anania. As a result, Pronomis Munroe & Mutuura, 1968 syn. n., Tenerobotys Munroe & Mutuura, 1971 syn. n., and Udonomeiga Mutuura, 1954 syn. n. are synonymized with Anania. The species formerly tretaed in Pronomis are transferred to Anania: Anania delicatalis (South, 1901) (Pyrausta) comb. n., Anania flavicolor Munroe & Mutuura, 1968 (Pronomis) comb. n., Anania profusalis (Warren, 1896) (Opsibotys) comb. n. The species and subspecies formerly treated in Tenerobotys are transferred to Anania: Anania subfumalis Munroe & Mutuura, 1971 (Tenerobotys) comb. n., Anania subfumalis continentalis (Munroe & Mutuura, 1971) (Tenerobotys) comb. n., Anania teneralis (Caradja, 1939) (Hapalia) comb. n., and Anania teneralis tsinlingalis (Munroe & Mutuura, 1971) (Tenerobotys) comb. n. Anania vicinalis (South, 1901) comb. n. (Pyrausta) is transferred from Udonomeiga to Anania. The apomorphic characters of Anania are also shared by the afrotropic Ethiobotys Maes, 1997, syn. n., and the species formerly treated therein are transferred to Anania: Anania amaniensis (Maes, 1997) comb. n., Anania ankolae (Maes, 1997) comb. n., Anania bryalis (Hampson, 1918) (Lamprosema) comb. n., Anania camerounensis (Maes, 1997) comb. n., Anania elutalis (Kenrick, 1917) (Pyrausta) comb. n., Anania epipaschialis (Hampson, 1912) (Nacoleia) comb. n., Anania lippensi (Maes, 1997) comb. n., and Anania ruwenzoriensis (Maes, 1997) comb. n. In contrast, Crypsiptya Meyrick, 1894 stat. rev. is reinstated as a valid taxon,