Habitat characteristics affecting use of lowland agricultural grassland by birds in winter

Almost a third of the bird species designated as Species of European Conservation Concern exploit agricultural grasslands, yet few studies have focused on their use as foraging habitats for birds. This study investigated the influence of variation in sward structure, grassland management and landscape variables on the use of 77 grass fields by 14 field-feeding bird species wintering on lowland mixed farmland in southern England. Multiple logistic regression was used to model the proportion of bird-count visits in which each species was encountered as a function of the recorded habitat variables for each field. Variation in sward height and density were associated with frequency of occurrence for 12 bird species and larger areas of bare earth and occurrence of winter grazing by stock animals were correlated with greater frequency of occurrence by 11 bird species. Two rapidly declining species, skylark ( Alauda arvensis) and yellowhammer ( Emberiza citrinella), were recorded more frequently on fields with higher numbers of seeding grasses. We suggest that mosaics of fields managed as short-term leys and permanent pastures with low-intensity cattle grazing over the autumn and winter would provide the combination of heterogeneous sward structure, areas of bare earth and presence of some seeding plants necessary to maximise the range of bird species able to use a given area of agriculturally improved grassland throughout the winter.