Four new species of Scarabapolipus (Acari: Podapolipidae) ectoparasites of scarabaeidae (Coleoptera) from the Western Pacific region

Four new species of the genus Scarabapolipus, ectoparasites of scarab beetles, are described from the Western Pacific region: S. glycyphanae n. sp. from Glycyphana fulvistemma Motschulsky collected in western Japan and from Glycyphana papua (Wallace) collected in northwestern Papua; S. nagaii and S. longichelae n. spp. from Protaetia ciocolatina (Wallace) collected in central Sulawesi and from Protaetia celebica (Wallace) in northern Sulawesi, Indonesia; and S. bisetosus n. sp. from Protaetia fusca (Herbst) collected in southeastern New Guinea. The new species are compared with the only known species in this genus, S. uchikawaii Husband & Kurosa, 1993, from the Japanese scarab beetle, Protaetia orientalis submarmorea (Burmeister), and with four other genera of podapolipid mites that are parasites of scarab beetles.