Mites (Acarina) associated with species of Trox (Coleoptera: Scarabaeidae)

Twelve mite species [Macrocheles penicilliger (Berlese), Macrocheles sp., Uroseius infirmus (Berlese), Uroseius sp., Uropoda orbicularis (Mueller), Cerratoma szekessyi Mahunka, Bakerdania decumana Krczal, Schwiebea nova (Oudemans), Schwiebea sp., Cisellipsopus microsporus Fain & Mahunka, Rhopalanoetus sp. and Listrophorus sp.] representing six families (Macrochelidae, Uropodidae, Pygmephoridae, Acaridae, Histiostomatidae and Listrophoridae) were found to be phoretic on four species of the scarabaeid genus Trox F. [T. sabulosus (L.), T. scaber (L.), T. hispidus (Pontoppidan) and T. cadaverinus Illiger]. Phoresy was observed for the first time in the following mite species: Listrophorus sp., Cisellipsopus microsporus, Bakerdania decumana and Uroseius infirmus. Uropoda orbicularis and Schwiebea nova are identified here as phoretic upon species of Trox for the first time.