Altitudinal and longitudinal variations in Pinus brutia (Ten.) of Crete Island, Greece: some needle, cone and seed traits under natural habitats

Four populations representing the entire natural distribution range of Pinus brutia (Ten.) in Crete (Greece) were sampled to determine: (1) Altitudinal variation within populations of Lassithi, Vorizia, Koustagerako, and Samaria, and; (2) Longitudinal variation among the four populations, based on morphological and anatomical differences in cone, needle and seed characteristics. Samples were collected from 10 trees per altitude, at three altitudes in each population. A total of 12 morphological and anatomical traits from needles, cones and seeds were analyzed to assess the altitudinal and longitudinal variations of the species in the island. The number of teeth per cm showed significant (P