New male reproductive organs of gymnosperms Permotheca colovratica sp. nov. from the Lower Permian of the Ural Mountains

A new species of male gymnosperm reproductive organs, Permotheca colovratica sp. nov. from the upper part of the Kungurian Stage (Lower Permian, Irenian Horizon, Koshelevskian Formation) of the Middle Cis-Urals and from the Kazanian Stage of the southern Cis-Urals and Volga River Basin, is described. These organs are rosette-like synangiate aggregations of microsporangia consisting of four sporangia fused at their bases. The adaxial surface of the synangia has a disc of attachment. The in situ pollen grains are of Falcisporites type. The morphologically similar genera Falcisporites Leshik emend. Klaus, Alisporites Daugherty emend. Nilsson, and Vesicaspora Shemel emend. Wilson et Venkatachala are analyzed. The morphogenetic trend in the evolution of the androphores Permotheca is discussed.