First record of the brachiopod Erymnaria in the Chruteren Member (Euthal Formation) from a new Palaeogene site in the Brülisau Schuppenzone of northeastern Switzerland (Canton St. Gallen) with remarks on shell asymmetry

A small, smooth-shelled rhynchonellid brachiopod, Erymnaria Cooper, 1959 was discovered in a previously unknown locality in the so-called Brülisau Schuppenzone (imbricate zone) of the South Helvetic region of northeastern Switzerland. It is the first record of this genus in this region and in Switzerland in general. It is comparable to the type species of Erymnaria , E. polymorpha (Massalongo, 1850). Its specific identity is currently undetermined, and the species is referred to Erymnaria sp. 1. The main characteristic of Erymnaria sp.1 is its asymmetric shell. This prompted a comparison of E . sp. 1 with other known asymmetric brachiopods. Questions of asymmetry and variability within the same genus and with brachiopods in general are discussed. Another, larger brachiopod species found at the new site is tentatively named Erymnaria ? sp. 2. The site can be assigned to the Chruteren Member (Euthal Formation) of Early Ypresian age, dated by large Foraminifera. Thus, the previous distribution area of the Chruteren Member can be extended.