Plant composition and diversity at edges in a semi-natural forest–grassland mosaic

As key components of landscapes, edges have received considerable scientific attention in anthropogenic ecosystems. However, edges in natural and semi-natural forest–grassland mosaics have received less attention, despite the fact that they cover a considerable proportion of these mosaic ecosystems. We studied forest edges in a semi-natural forest– grassland mosaic ecosystem of the Samobor Mountains (Croatia). Our aim was to compare the species composition, diversity and ecological indicator values of forest edges to those of the interior parts of the adjacent forest and grassland habitats. The vegetation was studied in 80 plots established in forest patch interiors, north-facing forest edges, south-facing forest edges and grassland interiors. We found that edges had a unique species composition, containing species from both the forest and the grassland interiors plus their own edge-related species (i.e. species that significantly preferred the edge habitat). These local edgerelated species did not correspond to regionallyidentified edge-related species. Compared to the forest and the grassland interiors, we revealed increased species richness in north-facing edges but not in southfacing edges. The mean light availability and nutrient supply indicator values of the edges were intermediate between those of the forest interiors and the grasslands. The mean soil moisture indicator values of the edges were similar to those of the grasslands. Our results show that edges form a unique component of forest–grassland mosaics, and they contribute considerably to landscape complexity, which should be taken into account during conservation decisions and habitat management.