Stratigraphic relations and habitat of the Bende fish teeth, southeastern Nigeria

The age and habitat of the Bende fish fossils have remained a matter of controversy with interpretations ranging between the Paleocene Imo Formation and the Eocene Ameki Formation. Outcrop-based sequence stratigraphic analysis presented in this work shows that the Imo and Ameki Formations together include three laterally extensive estuarine central basin/offshore mud rock units that are associated with the 59.7, 58.1, and the 56.8 ma maximum flooding surfaces, respectively. Field evidence and stratigraphic analysis show that the Bende fossil bones are hosted by the mud rock unit containing the 58.1 ma maximum flooding surface, while the Ameke bone fossils are contained within the condensed section associated with the 56.8 ma maximum flooding surface. Apart from highlighting the relevance of sequence stratigraphy in paleobiological interpretations, this study shows that the Bende fish fauna which Arua first assigned to the Eocene Ameki Formation indeed belongs to the Paleocene Imo Formation.