Development patterns in two hemlock-hardwood stands in southern New England

Two forest stands, composed primarily of northern red oak (Quercusrubra L.), red maple (Acerrubrum L.), and eastern hemlock (Tsugacanadensis (L.) Carr.), were studied by stand-reconstruction techniques to determine the pattern of development of canopy structure. One stand had originated following clear-cutting 87 years ago; the other, following catastrophic windthrow 44 years ago. Juvenile height growth of the hardwood species was much greater than that of hemlock and a stratified canopy developed by age 30 years, with hardwoods forming an overstory canopy above hemlock. Hemlocks maintained overstory positions only if they were 3 m or more in height immediately following canopy disturbance. In the older stand, hardwood height growth was about twice that of the tallest understory hemlocks during the first 30 years. The hardwood overstory slowed after that and grew at the same rate as the tallest understory hemlocks, which maintained a constant rate of height growth, and a constant to accelerating rate of basal area growth for much of the 87-year measurement period. The height growth of the tallest understory hemlocks was apparently limited in part by breakage of terminal shoots, caused by abrasion against branches of overstory hardwood crowns.