Devices, Methods, and Graphical User Interfaces for Navigating, Displaying, and Editing Media Items with Multiple Display Modes

An electronic device having a display and a touch-sensitive surface displays a mediate item representation corresponding to a sequence of images in a first display mode of user-selectable display modes, where the media item is part of a collection of ordered media items such that prior media item(s) precede the first media item and subsequent media item(s) follow the first media item in the order. In response to detecting touch input including movement in a direction, the device: upon determining that a subsequent media item corresponding to a second sequence of images is designated as being displayed in the first display mode, displays the subsequent media item in the first display mode, and upon determining that the subsequent media item is designated as being displayed in a second display mode different from the first display mode, displays the subsequent media item in the second display mode.