Image sensors having capacitively coupled pixels

An image sensor pixel may include a photodiode, a floating diffusion, and a transfer gate. Column readout circuitry coupled to the image sensor pixel via a column line. The image sensor pixel may have a pixel output path that is capacitive coupled to the column line via a capacitor. An input terminal of the capacitor at the image sensor pixel may be coupled to a pre-charging transistor. The pre-charging transistor may connect the input terminal of the capacitor to a grounding voltage. During readout operations, the pre-charging transistor may be activated before a row select transistor is activated to read out reset or image level signals. By capacitively coupling the image sensor pixel to the column line, pixel signal readout operations such as signal readout speed may be improved while pixel power may be reduced.