The mammal localities of Grand Daoui Quarries, Ouled Abdoun Basin, Morocco, Ypresian : A first survey

The unexpected discovery of mammals such as the earliest known proboscidean Phosphatherium escuilliei in the Ouled Abdoun phosphate Basin (Morocco), which is otherwise renowned for its very rich marine vertebrate fauna, has opened major new perspectives on the early African placental fauna. It was the impetus for several field parties by us between 1997 and 2001 which has allowed this survey of the geological, stratigraphic and paleontological context of Phosphatherium localities. This is the first important paleontological field work dealing with higher vertebrates in the Ouled Abdoun Basin since the work of C. Arambourg. Most of the material of Phosphatherium and other Ouled Abdoun mammals was found by local people searching for fossils for dealing, in an area where the economy is widely dominated by the mining development, in addition to more traditional but very poor agricultural resources. All Phosphatherium remains were found in the northeasternmost quarries of the Ouled Abdoun Basin, from a restricted quarrying area called Grand Daoui (fig. 3). In all Grand Daoui recognized localities (8), the mammals come from the lowermost Eocene level. The stratigraphically re-situated mammalian material was found in a bone-bed of the level called “Intercalaire Couches II/I” which is dated as early(iest) Ypresian on the basis of its relative stratigraphic position and its selachian fauna. At present, the occurrence of several mammal horizons in the “Intercalaire Couches II/I” (and Couche I ?) cannot be excluded, but their age difference seems not significant according to both the associated selachian taxa and the mammalian material, in present data. The bone-bed is very rich in macro-vertebrates. It includes especially large teeth of the shark Otodus obliquus which are the subject of very active research for dealing, and which explains the recent discovery of mammals such as Phosphatherium The level “Intercalaire Couches II/I” is made of a hardened phosphatic bioclastic limestone which is separated from underlying Paleocene levels by a major discontinuity. It is the first deposit of the Eocene sequence (megasequence C) in the Ouled Abdoun phosphate series. Deposition of this new sequence partly reworked the underlying Thanetian levels in the lowermost horizons of the level “Intercalaire Couches II/I”, such as the mammals bearing bone-bed. This explains the recovery of Thanetian selachians in the matrix of the holotype of P. escuilliei and its initial allocation t