Antropoėkologija Severo-Vostočnij Azii Čukotka, Kamčatka, Komandorskie ostrova

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adam_text СОДЕРЖАНИЕ Введение ....................................................................9 Глава 1 Некоторые итоги предшествующих популяционно-антропологических исследований в Северо-Восточной Азии ............................................... 2О Глава 2 Природные особенности Северо-Востока Азии ............................. зд Глава 3 Антропологический состав коренных народов Чукотки и Камчатки ........... φ Глава 4 Алеуты Командорских островов (соматологические наблюдения) ............. 8g Глава 5 Материалы по краниологии науканских эскимосов (к дифференциации арктической расы) .................................. log Глава 6 Дерматоглифика коренного населения Чукотки, Камчатки и Командорских островов ..................................... 1 19 Глава 7 Характеристика зубной системы населения северо-восточной Азии ..........130 Глава 8 Эритроцитарные системы крови в популяциях коренного населения Чукотки и Камчатки ................................ ι6ι 6 АНТРОПОЭКОЛОГИЯ СЕВЕРО-ВОСТОЧНОЙ АЗИИ Глава 9 Генетическое разнообразие населения Северо-Восточной Азии и этногенетические процессы в регионе Берингии и сопредельных территориях ................ 2о8 Глава 10 Морфологические особенности тела народов Северо-Восточной Азии ....................................238 Глава 11 Физиологический статус популяций Северо-Восточной Азии ................270 Глава 12 Физиологический статус популяций Северо-Восточной Азии ................296 Глава 13 Антропоэкологические связи на территории Северо-Восточной Азии .................................. gig Заключение Некоторые вопросы этногенеза и адапатции к окружающей среде .......................................323 Anthropology of North-Eastern Asia. Chukotka, Kamchatka, Commandor Islands ................................328 Литература ................................................................ Приложение Антропологические портреты отдельных представителей народов Чукотки и Камчатки .............................357 CONTENTS Introduction ........................................................... g Chapter 1 Some results of the population-anthropological researches in North-Eastern Asia ...........................................20 Chapter 2 Environmental specifics of Asia s North-East .................................29 Chapter3 Anthropological structure of the aboriginal populations of North-Eastern Asia and some problems of their ethnic genesis .................56 Chapter 4 The Aleuts of the Commander Islands (Somatological observations) ..............89 Chapter 5 Materials for the craniology of the Naukan Eskimos (Concerning subdivision of the Arctic race) .................................109 Chapter 6 Dermatogliphic traits of the aboriginal population of Chukotka, Kamchatka, and the Commander Islands ........................119 Chapter 7 Characteristic of dental system of the population of North-Eastem Asia ...........130 Chapter 8 Erythrocitic blood systems in the groups of aboriginal population of Chukotka, Kamchatka and the Commander Islands ........................161 8 АНТРОПОЭКОЛОГИЯ СЕВЕРО-ВОСТОЧНОЙ АЗИИ Chapter 9 Genetic diversity of the North-Eastern human population and ethnogenetic processes in the Bering Sea region and neighboring territories ..............................................208 Chapter 10 Body morphological specifics ofthe Koryak peoples of North-Eastern Asia ........238 Chapter 11 Physiological status of populations of North-Eastern Asia ......................270 Chapter 12 Age indications ofthe osteomorphic status and skeletal mineralization of hand among the aboriginal population of North-Eastern regions of Russia ........................................296 Chapter 13 Anthropo-ecological relationships in the territory of North-Eastern Asia ..........313 Conclusions ..........................................................323 Summary Bibliography Appendix Photo portraits ofthe representatives ofthe studied peoples of North-Eastern Asia ANTHROPOLOGY OF NORTH-EASTERN ASIA. CHUKOTKA, KAMCHATKA, COMMANDOR ISLANDS INTRODUCTION The expedition organized by the Institute of Ethnography of the USSR Academy of Sciences, and the Scientific Research Institute and Museum of Anthropology of Moscow State University carried out studies of the aboriginal population of Chukotka, Kamchatka and the Commander Islands from 1970 to 1982. The investigation program included racial characteristic, i.e. head measurement and description of facial soft tissues structure; body measurement and description, including adipose folds measurement in different parts of the body; taking teeth casts; taking palm and finger-prints. Besides the morphological characteristics, physiological ones were also studied, including determination of gaseous metabolism, and protein, lipid, and mineral metabolism. These indications basic ones for evaluation of adaptive characteristics of population. The work presupposed also determination of genetic markers, taking blood samples for laboratory physiological and genetic studies, as well as determination of gustatory sensitivity to PTC. The age indications of skeletal maturity and skeletal mineralization were investigated with application of the methods of radiophotography. Within the framework of various programs over 2000 individuals were investigated, both adults and non-adults, representing the Eskimos, the Chukchi, the Koryaks, the Itelmen, the Evens, and the Aleuts. CHAPTER 1. SOME RESULTS OF THE POPULATION-ANTHROPOLOGICAL RESEARCHES IN NORTH-EASTERN ASIA. The chapter discusses the results of the studies of aboriginal peoples carried out some time ago in North-Eastern Asia. The following results are of special significance. On the Asian littoral continuity was established between the population groups that had left the Uelen and Ekven cemeteries and the modern population (G.E Debets, M.G. Levin, V.P. Alekseyev). Genetic study of Siberian peoples and those living in North-Eastern Asia allows us to highlight the problems related to the evolution stability and dynamics of populations, as well as to genetic adaptation. The obtained results are evaluated against the background of the general picture of the population genetic processes in Siberia (Yu.G. Rychkov, V.A. Sheremet eva). Genetically, the Asian Eskimos do not form a separate group, which is confirmed by somatological and craniological analyses. The Chukchi and the Eskimos may be considered regular bearers of the Arctic complex of indications, which contradicts the Eskimo wedge hypothesis. In the aboriginal population s morphological-functional complex of indications we can reveal the features related to adaptation to the extreme environmental conditions of Arctic. CHAPTER 2. ENVIRONMENTAL SPECIFICS OF ASIA S NORTH-EAST. Environmental specifics of the Far East are determined by the region s geographic position, atmosphere circulation within the continent - ocean system, in winter - the influence exerted by ANTHROPOLOGY OF NORTH-EASTERN ASIA the Siberian anticyclone, adjacent cold-water seas, and the history of the region s development during protracted geological time. The majority of settlements in Chukotka and the Koryak plateau are situated in arctic and subarctic landscapes typified by extremely low environmental potential and extremely unfavourable habitation conditions characterized by strong deficiency of warmth, UV radiation (its deficiency is registered for 8 months in a year), and generally stern climatic conditions. Cold season lasts here for over 200 days, low temperatures (with absolute minimum under — 45 С о) is combined with strong winds, their maximal speed being over 40 m per second, more than 60 days of them with snowstorms. The index of weather harshness often exceeds 4.5 — 5 units (according to Bodman). The settlements located in the areas situated farther southward (in Kamchatka and the Commander Islands) are disposed in the boreal meadow-forest landscapes, and also enter the group with low environmental potential and unfavourable living conditions. Anyway, relatively favourable period in these regions is somewhat longer, index of sun radiation is higher, while deficiency of UV radiation is registered in the middle of the winter season only. The biogeochemical parameters are also different. The tundra landscapes are characterized by sharp deficiency of macro— and trace elements, which contributes to the emergence of biogeochemical deficiency endemics. CHAPTER 3. ANTHROPOLOGICAL STRUCTURE OF THE ABORIGINAL POPULATIONS OF NORTH-EASTERN ASIA AND SOME PROBLEMS OF THEIR ETHNIC GENESIS. Application of canonic analysis for determination of the position occupied by the peoples of North-Eastern Asia produces two clusters in the correlation field: one comprises the Eskimos, the Chukchi, the Itelmen, the Aleuts, the Koryaks and the Evens; another one — the peoples of the Altai- Sayan highland, the Yakuts and the Mongols. This clearly separates representatives of the Arctic race, on the one hand, and those of the Central Asian race, on the other hand. As for the Evens, they are related by their origin to the inland regions of Siberia, and hence are expected to be representatives of the Central Asian race, but they enter the North-Easter cluster. Evidently, a protracted stay in the Kamchatka Peninsula, amidst its aboriginal peoples accompanied by metisation with the local groups led to the situation when the Evens had acquired the features relating them to the Arctic race. It should be noted, however, that separate ethnic groups preserve to a certain extent their anthropological specifics. In connection with the above a rather polymorphic character of the Arctic race may be supposed. Unlike this race, the representatives of the Central Asian race look more homogeneous. Anyway, the said concerns the Altai and Tuva peoples, the Yakuts and the Mongols. CHAPTER 4. THE ALEUTS OF THE COMMANDER ISLANDS (SOMATOLOGICAL OBSERVATIONS). The chapter is based on the study of the Aleuts, aboriginal population of the Commander Islands, and of their metises with Russian people. The program of investigation included different head and facial measurements, as well as some so-called descriptive features commonly used to discriminate between the Caucasoid and Mongoloid races. Morphological traits of the Aleuts, even taking into account the fact that they developed with participation of definite Caucasoid admixture, form a combination that differs substantially from the Eskimo one. It should be pointed out that the contacts with the southern populations maintained along the Pacific coast played a considerable role in formation of this combination. CHAPTER 5. MATERIALS FOR THE CRANIOLOGY OF THE NAUKAN ESKIMOS (CONCERNING SUBDIVISION OF THE ARCTIC RACE). Craniological materials obtained from an Eskimo burial ground investigated in Naukan settlement are described. The results of their study are compared with craniological data on the Eskimos of Alaska, Canada and Greenland. The Eskimo craniological complex is substantially differentiated, much more than the populations of many areas in Eurasia. Comparison between a totality of features typical of the western and eastern Eskimo groups has confirmed the existence of western and eastern variant within the Eskimo complex as a whole. антропоэкология северо-восточной Азии CHAPTER б. DERMATOGLIPHIC TRAITS OF THE ABORIGINAL POPULATION OF CHUKOTKA, KAMCHATKA, AND THE COMMANDER ISLANDS. Dermatogliphic traits of 18 ethno-territorial aboriginal groups of North-Eastern Asia have been analyzed with application of the method of H. Cummins and Ch. Midlo: three Eskimo and seven Chukchi samples, as well as the Itelmen, the Evens and the Aleuts (one sample from each group). Totally 2221 individuals have been studied: 939 men and 1282 women. Men and women groups were analyzed separately. Multidimensional analysis was performed with the use of two independent methods (i.e. inter-group scale method and that of principal components). All the population of Chukotka and Kamchatka studied belong to the North-Asiatic Mongoloids. The next level of similarity brings the Eskimo, the Koryaks and the Itelmen closer to the southern and eastern Mongoloids. Differences between the local populations and ethnic groups of the region are higher than the general Siberian level. Within this range of diversity the following variants are distinguished: the Eskimo-Chukchi, the KoryakTtelmen and the Aleuts. The Chukchi and, especially, the Eskimo show the features of similarity with the southern Mongoloids. In the structure of the Koryaks, the Itelmen and, to for the most part, of the Evens of Kamchatka there is a significant south-Caucasoid admixture, which allows us to view Kamchatka as a specific race-breeding focus, without counterparts in the territory of Siberia. The limited size of the Aleutian sample possibly resulted in its sexual dimorphism and discrepancy of the results obtained from the comparative analysis of the Aleut men and women, which does not give grounds to draw any definite conclusions on the established specifics of the Aleuts. CHAPTER 7. CHARACTERISTIC OF DENTAL SYSTEM OF THE POPULATION OF NORTH-EASTERN ASIA. All studied groups according to the majority of odontologie indications are considered the representatives of the Eastern odontologie tree (all indications except for the distal crest of trigonide, lpa and Carabelli tubercle). Western odontologie indications are registered in all groups to the same degree except the Nunyamo, Lorino, Uelen and Kamchatka Chukchi, or slightly less than the Eastern ones. Noteworthy, this correlation is characteristic also of the ancient Eskimos of the Bering Sea region. The Eskimos of the late XIX — early XX cc. showed Eastern indications more clearly expressed, as compared to Western ones. The studied materials give grounds to single out certain Chukchi-Koryak group, uniting the northern, the Sireniki, and the Uelen Chukchi, and the Penzhinskaya Bay, the Coastal, the Palana, and Karaga Koryaks. This group according to the aggregate of indications both of Eastern and Western odontologie tree is close to the Greenland Eskimos, the Old Harbor Aleuts, and the Ulchi, as well as generalized groups of the Chukchi, the Koryaks, and the eastern Chukchi. The Itelmen occupy an isolated position in a generalized comparison of the groups; they show strong development both of Eastern and Western indications. The Evens stay most closely to the Aleuts of the Commander Islands; they show the both complexes developed to the same degree. They are most similar to the Uelen sample of ancient Eskimos, and the Chukchi as a whole. CHAPTER 8. ERYTHROCITIC BLOOD SYSTEMS IN THE GROUPS OF ABORIGINAL POPULATION OF CHUKOTKA, KAMCHATKA AND THE COMMANDER ISLANDS. Several populations of Chukotka and Kamchatka (the Chukchi, the Eskimos, the Koryaks, the Evens, the Itelmen, the Aleuts) were investigated during the expeditions of 1970-1982 supervised by V.P. Alekseyev, T.I. Alekseyeva and T.S. Balueva. The chapter discusses the genetic geography of the populations attributed to the following groups of blood: ABO, MN, Le, Ρ и Rh. The results have generally confirmed the well-known results obtained with the help of classic methods of morphological anthropology. According to genetic markers (and to morphology as well) the groups of Asian Eskimos and the Coastal Chukchi appeared to be most closely related to each other, on the one hand, and to the nomadic Chukchi, the Koryaks and the Evens, on the other hand. The Aleuts, the American Eskimos and rarely the Itelmen were the most distant from the clusters of other ethnic groups on the corresponding Tree Diagrams. The Koryaks are the most ANTHROPOLOGY OF NORTH-EASTERN ASIA 331 heterogeneous according to genetic frequencies of investigated blood systems. With the help of ABO blood system, vhich turns to be the most informative, as compared to the other mentioned above blood systems, it became possible to discriminate between the chronological stages of the Eskimos spreading in Arctic. The earliest stage was characterized by high frequency of r gene and low frequencies of p и q genes. The high frequency of p gene is most characteristic of the next stage, which is not typical of the present arctic populations in Asia. The latest stage of the Eskimos spreading shows relatively high East-Asian gene q frequency, which points to the relationship of these populations with those of the East Asiatic regions situated farther to the south. This feature is characteristic only of few populations of the Eskimos and of the Koryaks as a whole. Further analytical study is desirable. CHAPTER 9. GENETIC DIVERSITY OF THE NORTH-EASTERN HUMAN POPULATION AND ETHNOGENETIC PROCESSES IN THE BERING SEA REGION AND NEIGHBORING TERRITORIES. Complex information concerning genetic structure of the North-Eastern aboriginal population groups are discussed in the chapter. The history of population genetic studies of the Udeghe, the Nivkhs, the Evens, the Itelmen, the Koryaks, the Chukchi, the Eskimos and the Aleuts is presented and compared with adjacent large anthropological groups. The genetic connections among the studied ethnic groups were confirmed by serological, genetic-biochemical and molecular- genetic data. Genetic similarity of the Arctic anthropological type and the other Siberian population groups is established. The comparative analysis shows different taxonomical value both of individual genetic systems and their complexes for discussing the problems of historical genetics. The most abundant information about ethnic-anthropological value of the analyzed data may be derived from mtDNA, Y-chromosome and Gm haplotypes. The Eskimos wedge theory is discussed with the regard to population migrations from Asia to America. Spatial vector of the gene frequencies reflects a series of migrations directed from the south to the north along the Asian coast. This study also confirms existence of genetic connections between the studied North-Eastern ethnic groups and Siberian and Central Asian populations. CHAPTER 10. BODY MORPHOLOGICAL SPECIFICS OF THE KORYAK PEOPLES OF NORTH-EASTERN ASIA. The insignificant difference observed in the somatotype of North-Eastern Asia population evidently can be explained both by influence of different climatic conditions (for instance, the Kamchatka and Chukotka peoples) and certain geographic isolation. Metisation with the migrant groups must have played its role as well, as it can be seen from the relationships between the Itelmen and the Evens. Finally, we must take into account some morphological features in common that evidence earliest single origin of the peoples of North-Eastern Asia (the Aleuts morphology can serve as an example). This insignificant morphological differentiation apparently developed within the Arctic race, while the morphological features typical of the representatives of the Central Asian race are traced as an admixture only that occurred as a result of metisation of the aboriginal and migrant population. CHAPTER 11. PHYSIOLOGICAL STATUS OF POPULATIONS OF NORTH-EASTERN ASIA. The stability and variability of physiological status was investigated in several populations of North-Eastern Asia (the Kamchadals — recent migrants to Kamchatka, the Coastal Chukchi and the Eskimos). The problems of stability and variability of physiological status of the local population are discussed from the viewpoint of the concept of physiological homeostasis. The physiological status of the Kamchadals and the migrants shows general similarity. The difference between the physiological status of the Eskimos and the Chukchi is significant. The violation of physiological homeostasis and a number of medical and sociological characteristics indicate a decrease of the Eskimos adaptability to the environment. АНТРОПОЭКОЛОГИЯ СЕВЕРО-ВОСТОЧНОЙ АЗИИ CHAPTER 12. AGE INDICATIONS OF THE OSTEOMORPHIC STATUS AND SKELETAL MINERALIZATION OF HAND AMONG THE ABORIGINAL POPULATION OF NORTH-EASTERN REGIONS OF RUSSIA. The ossification processes were studied on the Chukchi-Eskimos and the Eveno-Koryak children from the Chukotka Peninsula and Kamchatka respectively. The researches were carried out in the early 1970-s. The results showed that the pace of skeletal development of the discussed groups is similar to standard (method TW-2). The reveled pace is faster, as compared to those established in a series of the other rural aboriginal populations. The obtained results allow supposing the influence of moderate stress factors of social nature. The retrospective study of the menarche age confirms the supposition on advanced biological development in the region under study, which may be related to the transformation of traditional culture and life-style that took place in the territory. The obtained data are closely associated with the specifics of age dynamics observed on hand bones of the adult population in the studied regions. In particular, a high (in inter-group aspect) pace of aging process is registered in a number of aboriginal population groups of Kamchatka and Chukotka. CHAPTER 13. ANTHROPO-ECOLOGICAL RELATIONSHIPS IN THE TERRITORY OF NORTH-EASTERN ASIA. The populations of North-Eastern Asia live in the territories differentiated mostly by the level of favourable, or less favourable influence on human organism. Geographically, the ecological conditions clearly become more favourable, moving from north to south — from Chukotka to Kamchatka. To analyse the anthropo-ecological relationships the geographical method was applied, widely used in anthropological studies for a long time. Judging from the absolute and relative characteristics of body size of the aboriginal peoples of North-Eastern Asia, all of them have high heat regulation properties and are typical representatives of the Arctic adaptive type. In Chukotka this type is represented by a more massive variant, while in Kamchatka — by a more miniature one. When discussing dominating natural factors of environment in the territories of Chukotka and Kamchatka that cause adaptive reactions in human populations, we should mention first of all the level of extreme characteristics, and basic reaction from human organism — growth of energetic characteristics developed both in morphology and physiology. CONCLUSION. SOME QUESTION OF ETHNIC GENESIS AND ADAPTATION TO THE ENVIRONMENT The project of studying the aboriginal population of North-Eastern Asia presented in this book logically continued the Jesup expedition of the early XX century, as well as the fundamental investigations on the anthropology of the Far East carried out by G.F. Debets and M.G. Levin. The conclusion on the single origin of the aboriginal peoples of Kamchatka and Chukotka, and their attribution to different variants of the Arctic race should be considered basic result of the researches undertaken by the national anthropologists before. This paved the way to the solution of the protracted debate between the adherents of the palaeoasiatic hypothesis of the origin of the North-Eastern Asian peoples, and those who supported the hypothesis of the Eskimo wedge. The problem was solved in favour of the former, claiming the single origin of the peoples inhabiting North-Eastern Asia. Studies of the aboriginal peoples of Chukotka, Kamchatka, and the Commander Islands were resumed m relation with significant extension of the methods applied in anthropological investigations, which allow suggesting much more comprehensive characteristics of the local population s physical specifics. The scope of problems investigated by physical anthropology was also extended substantially, now it includes the questions of human populations adaptation to the environment, modes of interaction with the environment, and the historical dynamics of this interaction. In recent investigations of special importance are researches of erythrocytic and serum blood systems producing equally reliable information both for establishment of population s genetic relationships and assessment of specific features of population s adaptation to the extreme habitation conditions. Methods of statistical processing anthropological materials are now much ANTHROPOLOGY OF NORTH-EASTERN ASIA 333 more advanced, which provides us with better possibility for objective evaluation of the investigation results. The results of our research work in North-Eastern Asia have been partly published, but not in entirety, so that the scholars participating in the program consider it necessary to resume the theme of the anthropology of the North-East in order to highlight in details the problems of ethnic genesis of the Arctic and subarctic peoples, and their interrelation with the environment. New anthropological data confirm attribution of the modern aboriginal population of Chukotka, Kamchatka, and the Commander Islands to the Arctic race. Variations registered within it are related by its origin to certain ethnic and geographic isolation some peoples lived in, in-population variability, and, probably, early admixture of Eastern and Southern Mongoloid elements, similar to what we know about the Aleuts. Application of R. Knussman and Malyutov-Pasekov methods of digital taxonomy (Алексеева и др., 1978) for establishing genetic distances between the peoples of Chukotka, Kamchatka, the Commander and the Aleut islands and Alaska clearly shows their geographic isolation. Two groups of islander Aleuts are singled out, the both are related to the Alaska Eskimos; the Itelmen occupy rather isolated position as one of the most ancient peoples in North-Eastern Asia, while the Chukchi and Eskimos are very close to each other; finally, the Koryaks are rather closely related to them. The genetic analysis perfectly correlates with the anthropological, historical and ethnological data. Representatives of the Arctic race differ from all continental Mongoloids by lesser development of the Mongoloid indications in the eye zone, and more dark-coloured skin. In this aspect they are closer to the Eastern and Southern Mongoloids than those inhabiting the inland regions of the continent. The results obtained by the recent investigations confirm the genetic unity of the peoples of Kamchatka, Chukotka, and the Commander Islands, and completely discard the hypothesis on the Eskimo wedge. The genetic unity of the peoples of North-Eastern Asia is convincingly evidenced by stable character of some genetic markers, in particular, 100% of individuals show rhesus-positive gene. From Chapter 8 it is clear that in 20 groups from 22 investigated ones the rate ipf rhesus-positive gene (D) reached 100%. In two groups only (the Chukchi of the Bering zone and the Karaga Koryaks) the admixture of rhesus-negative gene (d) makes up less than one tenth of percent. In the authors view, this situation was probably caused by the Caucasoid influence, since the both groups in question are coastal ones. The idea on the usage of rhesus-positive gene to prove the single genetic origin of the peoples of Chukotka, Kamchatka, and the Commander Islands was already suggested (Рычков, Шереметьева, 1972; Алексеев, 1974), but the studied materials were too limited then. The obtained genetic characteristics more clearly evidence specifics of ethnic history than environmental conditions. At the same time, 100% rhesus-positive indication may be regarded as a specific feature of adaptive character that prevents birth complications. The investigations carried out by Yu.G. Rychkov and V.A. Sheremet yeva among the peoples of Chukotka and Commander Islands demonstrate the rigid pressure exerted by natural selection. Judging from the dermatogliphic characteristics, population of Chukotka and Kamchatka enters the circle of the North Asiatic Mongoloids subdivided into several variants more closely related to one another. The first variant is represented by the Eskimos and Chukchi, the second one — by the Koryaks and Itelmen, the third variant includes the Aleuts. The Chukchi and Eskimos, the latter in particular, show certain similarities with the Southern Mongoloids. Among the Kamchatka Koryaks, Itelmen and Evens an admixture of Southern Caucasoid elements is registered, which allows the authors (see Chapter 6) regard Kamchatka as a specific focus of race formation process having no analogies in the territory of Siberia. Apparently, here we deal with an insignificant Caucasoid admixture related to the Russian Cossaks exploration of Kamchatka, formation of the local group of the Kamchadals, and their subsequent metisation with the aboriginal population. An insignificant Caucasoid admixture is also registered among the Kamchatka peoples according to the odontological data (see Chapter 7). In opinion of the author of this chapter N.A. Dubova, there are no grounds to relate this phenomenon to the late Caucasoid admixture, in particular, to the coming of the Russians, though the latter supposition must not be excluded. Similar situation was observed also among the peoples of the Amur basin, the Ainu, and some groups of Australians. Evidently, this problem needs further investigations. The local differentiation of the extreme North-East of Asia is established by the blood group factors as well (Chapter 8). The Chukchi and the Asian Eskimos show the highest indications of 334 АНТРОПОЭКОЛОГИЯ СЕВЕРО-ВОСТОЧНОЙ АЗИИ ________________ closeness; the Koryaks and Evens demonstrate rather similar relationship. The Aleuts and the American Eskimos are the least close from them, which well corresponds to the latter s geographical separation from the Asian populations. Within the Eskimo population certain closeness m physical appearance show the Asian and the Alaska Eskimos, while the Eskimos of Canada and Greenland are essentially different from the populations living in the Asian continent. It is highly evident that the anthropological differentiation is to a great extent determined by the distance factor, this observation being confirmed by the results of thesomatological investigations. , . The local anthropological differentiation of the modern peoples of North-Eastern Asia is confirmed by the analysis of craniological samples dating from the XIX-XX cc. and characterise all the Eskimo groups. The Asian Eskimos turned to be most heterogeneous, the least heterogeneous are the Greenland ones (Chapter 5). Among the factors that had caused this homogeneity might be growing influence of the natural selection during the settlement process, and, consequently, limited variability of indications. Long ago A. Hrdlička (1930) considered the Greenland variant as a result of the Arctic adaptation. He was the first to point out the role adaptation to the Arctic climate could have played in formation of the specific physical features of the Far North population. Concluding the section devoted to the problems of racial and ethnic genesis of the peoples of Chukotka, Kamchatka, and the Commander Islands it should be reasonable to dwell upon the researches in the field of genetic structure of the population of North-Eastern Asia (Chapter 9). The author of the chapter V.A. Spitsyn states that most valuable information for solving the problems of historical genetics may be derived from the data on haplotypes and haplogroups in the immune systems , mt DNA, and Y-chromosome. The character of gene frequency variability mirrors the consecutive, step-by-step pattern of the migrations that were directed from the South to the North along the Asian coast. We can see that the somatological researches revealing southern relationships of the peoples of North-Eastern Asia are seriously confirmed by the genetic data. Another series of problems is related to the anthropological studies of the North-Easţ Asian peoples, that is, to their morphophysiological characteristics that may be considered in the light of their adaptation to the extreme environmental conditions of the Far North. In the body structure of the aboriginal population of the Arctic and subarctic regions many features in common are observed, independently of the peoples origin. Relatively insignificant body length (according to the inter-group scale) is combined with relatively significant weight, as well as thorax, waist, and buttocks circumference, and strongly developed bone and muscle mass. These features are typical not only of the inhabitants of Chukotka, Kamchatka, and the Commander Islands, but of different territorial groups living in the American Arctic regions, as well as the forest Nentsy from the Pur River basin and the Lapps of the Kola Peninsula. When evaluating the morphological specifics of the peoples inhabiting North-Eastern Arctic, high concentration of the people of athletic constitution should be pointed out in the male selection (Chapter 10). In some groups this indication exceeds 50%. This proportion is much higher than what we know about other peoples living in the inland and western regions of Northern Asia. For instance, among the Buryats representatives of the athletic constitution make up only 15%. As for women, they show specific features related to the athletic constitution to a lesser extent, but in general the female selection also demonstrates high indications of physical development. Such phenomena as relatively high proportion of muscle constitution type in the male selection, and mesoplastic type in the female one are characteristic of all the groups living under the conditions of cold stress. We cannot exclude that this feature had emerged as a result of a long-term adaptation of human populations to the extreme environment, since strong development of muscle body mass intensifies basic metabolic process and thermal energy production. Probably, physical stresses had contributed to strong development of bone and muscle components in the body structure of the Arctic aboriginals. Thus, the individuals of athletic constitution among the aboriginal population of Chukotka whose traditional occupation was hunting sea animals is higher than that of the Kamchatka peoples traditionally engaged in reindeer-breeding, fishing and hunting. Certain age characteristics of the aboriginal inhabitants of the Asian North-East draw special attention: biologically, they are older than their officially registered age, with early manifestations of involution changes in the skeletal structure (Chapter^). If we interpret these indications as the signs of physical maturity, we shall acknowledge that the life cycle of the Arctic aboriginals is somewhat ANTHROPOLOGY OF NORTH-EASTERN ASIA 335 shortened. This supposition is confirmed by the fact of their insignificant life duration, though this circumstance might be determined also by the insufficient hygienic level and low social status of the Arctic peoples. Among the functional indications reflected in the body structure cylindrical shape of thorax should be mentioned, which is not typical of the inhabitants of the moderate climatic zone with high proportion of conical and flattened shapes. Under the conditions of increased basic metabolism and heat regulation high capacity of lungs is also considered an adaptive feature. Significant development of thorax in the Arctic conditions is related to hypoxia syndrome caused by deficiency of ascorbic acid and related to its increased expense in the intensified metabolic process (Неверова и др., 1972). The oxygen deficiency associated with the specific features of metabolism leads to intense blood producing related not only to increased of haemoglobin content, but also to strong development of the marrow zone. Radiological investigations of the Arctic people skeletons have shown their massive bones with wide marrow channels and thin compact layers. This feature is of key significance, because skeleton is a strong centre of haemoglobin synthesis, this respiratory protein guaranteeing processes of energy exchange in organism. The aboriginal inhabitants of Arctic demonstrate such characteristic physiological features as increased blood circulation, high rate of proteins and fats in blood serum, and their increased utilisation. There are sufficient data at our disposal to suppose the existence of inherited mechanisms regulating metabolic processes. These problems are discussed in details in the works dealing with the process of human populations adaptation within the territory of ecumene (Алексеева, 1977, 1986). In relation with the said it would be expedient to point out the investigation of basic metabolism among the inhabitants of the Circumpolar regions carried out by a joint team of researchers from the USA, Canada, and Russia (Leonard et al., 2002). The comparison of basic metabolism assessment among the aboriginal and migrant population shows statistically significant difference, the indication reaching 19% and 7% respectively. The authors are inclined to think that in the process of adaptation to cold climate both environmental and genetic factors play substantial role, in particular the thyroid gland hormones. In order to establish the chronology of formation of the anthropological specifics of the North- Eastern Asian population, and those of adaptive character in particular, it is necessary to consider the population of the ancient Bering Sea region known from the Uelen and Ekven cemeteries. Especially numerous are the craniological samples from Ekven. The chronological scope of the cemetery is very wide, covering the period from the 2nd с. ВС to the 14th с AD. G.E Debets was the first to investigate the skulls from these cemeteries, and he pointed to a specific structure of the Eskimo skulls: combination of very high cranial vault and extremely wide cranial base (Дебец, 1975). G.E Debets did not suggest any explanations of these features, but it is highly probable that they were related to the adaptive changes in cranium structure. The skull shape of the Eskimos buried in the Ekven and Uelen cemeteries practically does not differ from the modern one. The population is characterized by long and high head and very narrow nose. V.P. Alekseyev pointed out the adaptive nature of such skull shape in his publication concerning bipolar races (1985). Identical skull shape is typical of the population of Terra del Fuego. Possibly, this is related to some specific character of insolation, or some other geographical factors. Of special interest is the striking homogeneity of the Eskimo samples from the Ekven and Uelen cemeteries revealed by our investigations with the help of the Datascope system. During fifteen centuries of the cemeteries functioning the same anthropological type was reproduced without any visible changes. In my opinion, only one explanation can be suggested in this case: the natural selection under the extreme Arctic conditions was conducive to preservation of the anthropological type most adequately corresponding to the local conditions. The discussed type demonstrates such features as long head, very high skull, very large face and very narrow nose. The range of in-group indications variability is strongly reduced, especially in the female series. Such fact as reduction of in-group morphological variability in the human populations living in extreme habitation conditions was registered more than once, when studying modern population of various environmental niches. Consequently, we may state rather definitely that environment played an important role in formation of the ancient population of the Bering Sea coast. антропоэкология северо-восточной Азии Stability of the Eskimo anthropological type for almost two thousand years of their history was established long ago by M.G. Levin (1949). In the light of the recent investigations of the modern and ancient population of the Bering Sea zone this phenomenon can be explained by the rigid pressing exerted by the extreme Arctic conditions. This conclusion confirms the idea formulated by A. Hrdli ka concerning adaptive nature of the Greenland Eskimos skull structure, and the results obtained by Yu.G. Rychkov and V.A. Sheremet yeva, who studied the peoples of Chukotka and the Commander Islands, and explained their specific anthropological features by the pressure of natural selection (Шереметьева, Рынков, 1978). The morphological specifics of the body structure and certain features of metabolism established for the modern population are also associated with adaptation to the extreme conditions of the Far North. Basing on the morpho-physiological characteristics of the region s aboriginal population, the Arctic adaptive type was singled out and characterised as a standard biological response to the complex challenge of geographic conditions, with cold stress being most important (Алексеева, 1977). Geographically, the investigated region does not comprise strongly different environmental niches, unlike, say, Central Asia (Антропоэкология ..., 2005). The territories of Chukotka, Kamchatka, and the Commander Islands differ mostly by the level of favourable or unfavourable influence on human organism. The degree to what subsistence conditions may be considered favourable was assessed by a set of parameters including landscapes productiveness; chemical formulas of natural food resources, both of plant and animal origin; hydro-chemical properties of surface waters (Chapter 13). The more to the south a territory is situated, the more favourable its conditions are. The same gradient is revealed in absolute and relative body parameters of the aboriginal peoples of Chukotka and Kamchatka. All population groups show high thermorégulation mechanisms that demonstrate distinct reaction of organism to the influence of cold climate. At the same time, in the territory of Chukotka features of the Arctic adaptive type manifest themselves in a more massive variant, while in the territory of Kamchatka we find them in more miniature one. The massive variant is related to the extreme habitation conditions, and the miniature one — to more favourable environment. When discussing the influence of the factors of natural environment on human populations in the Far North- East of Asia, the level of extreme should be first pointed out as a leading factor, while basic reaction of human organism should be considered increased energy characteristics, which are achieved both by changes in body structure and metabolic processes.
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spellingShingle Antropoėkologija Severo-Vostočnij Azii Čukotka, Kamčatka, Komandorskie ostrova
Indigenes Volk (DE-588)4187207-1 gnd
Anthropologie (DE-588)4002230-4 gnd
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title Antropoėkologija Severo-Vostočnij Azii Čukotka, Kamčatka, Komandorskie ostrova
title_alt Anthropology of North-Eastern Asia. Chukotka, Kamchatka, Commandor Islands
title_auth Antropoėkologija Severo-Vostočnij Azii Čukotka, Kamčatka, Komandorskie ostrova
title_exact_search Antropoėkologija Severo-Vostočnij Azii Čukotka, Kamčatka, Komandorskie ostrova
title_full Antropoėkologija Severo-Vostočnij Azii Čukotka, Kamčatka, Komandorskie ostrova Naučno-Issledovatelʹskij Institut ... [Otv. red. T. I. Alekseeva ...]
title_fullStr Antropoėkologija Severo-Vostočnij Azii Čukotka, Kamčatka, Komandorskie ostrova Naučno-Issledovatelʹskij Institut ... [Otv. red. T. I. Alekseeva ...]
title_full_unstemmed Antropoėkologija Severo-Vostočnij Azii Čukotka, Kamčatka, Komandorskie ostrova Naučno-Issledovatelʹskij Institut ... [Otv. red. T. I. Alekseeva ...]
title_short Antropoėkologija Severo-Vostočnij Azii
title_sort antropoekologija severo vostocnij azii cukotka kamcatka komandorskie ostrova
title_sub Čukotka, Kamčatka, Komandorskie ostrova
topic Indigenes Volk (DE-588)4187207-1 gnd
Anthropologie (DE-588)4002230-4 gnd
topic_facet Indigenes Volk
Anthropologie
Ferner Osten Russland Nord
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