Revision of the “celia clade” of Pseudodebis Forster, 1964, with Two New Species and Notes on Papilio phorcys Fabricius, 1793 (Lepidoptera: Nymphalidae: Satyrinae)

The species-level classification of the “ celia clade” of the nymphalid butterfly genus Pseudodebis Forster, 1964, is revised as part of ongoing revisionary work on this genus. The “ celia clade” contains three species, of which two, Pseudodebis darrenthroopi Nakahara & Willmott, n. sp. and P. tigrillo Nakahara & Willmott, n. sp., are described and named herein based on morphology and molecular data. Consequently, we increase the described species diversity of Pseudodebis to 13, with a remarkable six species occurring in the trans-Andean region. We discuss five specific epithets associated with the clade and designate a neotype for Papilio celia Cramer, 1780, and lectotype for Taygetis keneza Butler, 1869, based on the same specimen, thus retaining the status of the former name as a senior objective synonym. We also provide a brief historical review for Papilio phorcys Fabricius, 1793, an enigmatic name currently synonymized under Papilio celia . Nevertheless, we were unable to locate the syntype(s) for this name and the identity of Papilio phorcys remains uncertain, so we retain the current synonymy as a parsimonious approach.