Habitat-related differences in the pre-copulatory guarding and copulation behavior of Neohelice granulata (Brachyura, Grapsoidea, Varunidae)

Mating behavior strategies of brachyuran crabs are flexible depending on the habitat and social context. We evaluate pre-copulatory guarding in Neohelice granulata and describe the unusual copulatory behavior. Laboratory experiments were conducted to test whether factors such as male size, presence...

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