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Subspecies and Distribution. Ep. pyrropus EF. Cuvier, 1833 — S Cameroon to Mayombe Forest in S Gabon and SW Republic of the Congo. E p. akka de Winton, 1895 — E DR Congo, Uganda, and W Rwanda. FE p. leonis Thomas, 1905 — Liberia and Sierra Leone. E p. leucostigma Temminck, 1853 — S Ghana. Ep. mandingo Thomas, 1903 — Gambia and S Senegal. F.p. nigrensis Thomas, 1909 — S Nigeria, between the Niger and Cross rivers. Ep. niveatus Thomas, 1923 — Ivory Coast. Ep. pembertoni Thomas, 1904 — N Angola and
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